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	RNF115/BCA2 E3 Ubiquitin Ligase Promotes Breast Cancer Cell Proliferation through Targeting p21(Waf1/Cip1) for Ubiquitin-Mediated Degradation
	Wang ZH1; Nie Z2; Chen WL3; Zhou ZM1; Kong QH1; Seth AK4; Liu R*1; Chen CS*1; liurong@mail.kiz.ac.cn; chenc@mail.kiz.ac.cn
	2013
发表期刊	NEOPLASIA
卷号	15期号:9页码:1028-1035
合作性质	其它
摘要	The E3 ubiquitin ligase RING finger protein 115 (RNF115), also known as breast cancer-associated gene 2 (BCA2), has previously been reported to be overexpressed in estrogen receptor alpha (ER alpha)-positive breast tumors and to promote breast cell proliferation; however, its mechanism is unknown. In this study, we demonstrated that silencing of BCA2 by small interfering RNAs (siRNAs) in two ER alpha-positive breast cancer cell lines, MCF-7 and T47D, decreases cell proliferation and increases the protein levels of the cyclin-dependent kinase inhibitor p21(Waf/Cip1). The protein stability of p21 was negatively regulated by BCA2. BCA2 directly interacts with p21 and promotes p21 ubiquitination and proteasomal degradation. Knockdown of p21 partially rescues the cell growth arrest induced by the BCA2 siRNA. These results suggest that BCA2 promotes ER alpha-positive breast cancer cell proliferation at least partially through downregulating the expression of p21.
资助者	This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.).
收录类别	SCI
语种	英语
资助者	This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.). ; This study was supported by National Key Basic Research Program of China (2013CB910900), National Natural Science Foundation of China (81072162, 81120108019, 81272930, and U1132605), Top Talents Program of Yunnan Province, China (2010CI114), Science and Technological Key Project of Yunnan Province (2012FB185), and “Western Light” Talents Training Program of Chinese Academy of Sciences (to Q.K.).
WOS记录号	WOS:000328134800004
引用统计
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/7771
专题	科研部门_肿瘤生物学（陈策实） 科研部门_动物模型与人类重大疾病机理重点实验室
通讯作者	liurong@mail.kiz.ac.cn; chenc@mail.kiz.ac.cn
作者单位	1.Key Laboratory of Animal Models and Human Disease Mechanisms of Chinese Academy of Sciences and Yunnan Province, Kunming Institute of Zoology, Kunming, Yunnan, China 2.Department of Neurology, The First Affiliated Hospital of Kunming Medical University, Kunming, Yunnan, China 3.Department of Breast, The Third Affiliated Hospital of Kunming Medical University, Kunming, Yunnan, China 4.Sunnybrook Health Sciences Centre, University of Toronto, Toronto, Ontario, Canada
推荐引用方式 GB/T 7714	Wang ZH,Nie Z,Chen WL,et al. RNF115/BCA2 E3 Ubiquitin Ligase Promotes Breast Cancer Cell Proliferation through Targeting p21(Waf1/Cip1) for Ubiquitin-Mediated Degradation[J]. NEOPLASIA,2013,15(9):1028-1035.
APA	Wang ZH.,Nie Z.,Chen WL.,Zhou ZM.,Kong QH.,...&chenc@mail.kiz.ac.cn.(2013).RNF115/BCA2 E3 Ubiquitin Ligase Promotes Breast Cancer Cell Proliferation through Targeting p21(Waf1/Cip1) for Ubiquitin-Mediated Degradation.NEOPLASIA,15(9),1028-1035.
MLA	Wang ZH,et al."RNF115/BCA2 E3 Ubiquitin Ligase Promotes Breast Cancer Cell Proliferation through Targeting p21(Waf1/Cip1) for Ubiquitin-Mediated Degradation".NEOPLASIA 15.9(2013):1028-1035.

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