Translocation of microbes and changes of immunocytes in the gut of rapid- and slow-progressor Chinese rhesus macaques infected with SIVmac239

	Translocation of microbes and changes of immunocytes in the gut of rapid- and slow-progressor Chinese rhesus macaques infected with SIVmac239
	Zhang LT 1,2; Tian RR 1,3; Xia HJ 1; Xia XS 2; Pang W1; Zheng YT1; Zheng HY 1,4; Pan GQ 5; Tuo XY 5; pangw@mail.kiz.ac.cn ; zhengyt@mail.kiz.ac.cn
	2016
发表期刊	IMMUNOLOGY
卷号	147 期号:4 页码:443-452
摘要	HIV/SIV infection can cause severe depletion of CD4+ T cells both in plasma and mucosa; it also results in damage to the gut mucosa barrier, which makes the condition more conducive to microbial translocation. In this study, we used simian immunodeficiency virus (SIV)-infected Chinese rhesus macaques to quantify the extent of microbial translocation and the function of immune cells in the entire gastrointestinal tract and to compare their differences between rapid and slow progressors. The results showed that in the slow progressors, microbial products translocated considerably and deeply into the lamina propria of gut; the tissue macrophages had no significant differences compared with the rapid progressors, but there was a slightly higher percentage of mucosal CD8+ T cells and a large amount of extracellular microbial products in the lamina propria of the intestinal mucosa of the slow progressors. The data suggested that although microbial translocation increased markedly, the mucosal macrophages and CD8+T cells were insufficient to clear the infiltrated microbes in the slow progressors. Also, therapies aimed at suppressing the translocation of microbial products in the mucosa could help delay the progression of SIV disease.
关键词	Mucosa Lamina Propria Microbial Translocation
资助者	This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02).
收录类别	SCI
语种	英语
资助者	This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02). ; This work was supported in part by grants from the National Basic Research Programme of China (2012CBA01305), the National Natural Science Founda- tion of China (81172876, 81471620, 81273251, U1202228, 81571606), the Key Scientific and Technological Pro- gramme of China (2012ZX10001-007, 2013ZX10001-002, 2014ZX10005-002-006), Natural Science Foundation of Yunnan Province (2014FB181) and the Knowledge Inno- vation Programme of the Chinese Academy of Sciences (KJZD-EW-L10-02).
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/9518
专题	科研部门_分子免疫药理学（郑永唐）科研部门_动物模型与人类重大疾病机理重点实验室
通讯作者	pangw@mail.kiz.ac.cn; zhengyt@mail.kiz.ac.cn
作者单位	1.Key Laboratory of Animal Models and Human Disease Mechanisms of Chinese Academy of Sciences & Yunnan Province, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, 650223, China 2.Faculty of Life Science and Technology, Kunming University of Science and Technology, Kunming, Yunnan, 650500, China 3.Kunming College of Life Science, University of Chinese Academy of Sciences, Kunming, Yunnan, 650204, China 4.School of Life Sciences, University of Science and Technology of China, Hefei, Anhui, 230026, China 5.The Pathology Department, First Affiliated Hospital of Kunming Medical University, Kunming, Yunnan, 650032, China
推荐引用方式 GB/T 7714	Zhang LT,Tian RR,Xia HJ,et al. Translocation of microbes and changes of immunocytes in the gut of rapid- and slow-progressor Chinese rhesus macaques infected with SIVmac239[J]. IMMUNOLOGY,2016,147(4):443-452.
APA	Zhang LT.,Tian RR.,Xia HJ.,Xia XS.,Pang W*.,...&zhengyt@mail.kiz.ac.cn.(2016).Translocation of microbes and changes of immunocytes in the gut of rapid- and slow-progressor Chinese rhesus macaques infected with SIVmac239.IMMUNOLOGY,147(4),443-452.
MLA	Zhang LT,et al."Translocation of microbes and changes of immunocytes in the gut of rapid- and slow-progressor Chinese rhesus macaques infected with SIVmac239".IMMUNOLOGY 147.4(2016):443-452.

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