Taxonomic revision of the Chinese Limnonectes (Anura, Dicroglossidae) with the description of a new species from China and Myanmar.

	Taxonomic revision of the Chinese Limnonectes (Anura, Dicroglossidae) with the description of a new species from China and Myanmar.
	Suwannapoom C 1,2; Yuan ZY 1,3; Wang LJ 7; Nguyen TS 8; Nguyen TQ 8; Murphy RW 1,9; Sullivan J 10; Mcleod DS 11; Chen JM 1,4; Che J*1,3; Hou M 5; Zhao HP 6; chej@mail.kiz.ac.cn
	2016
发表期刊	ZOOTAXA
卷号	4093 期号:2 页码:181-200
摘要	Phylogenetic reconstructions derived from DNA sequence data play a central role in documenting the number of species in a complex. Such analyses are pointing to the existence of many cryptic species, especially in poorly understood groups such as the genus Limnonectes, and the L. kuhlii species complex in particular. To understand the Limnonectes frogs of China, we reconstruct the major matrilineal genealogy of Limnonectes from China and Southeast Asia based on 12S rRNA, tRNA^Val and 16S rRNA gene sequences. Based on new data we recognize five species of Limnonectes in China including L. bannaensis, L. fujianensis, L. fragilis, L. taylori (new record), and a new species from southern China and Myanmar. Phylogenetically, the new species is more closely related to the clade comprising L. taylori, L. megastomias, L. isanensis, L. nguyenorum, and L. jarujini from Thailand than to other Chinese species. This study supports previous findings of sympatric members of a species complex that are not each other's closest relatives.
关键词	Indochina Limnonectes Longchuanensis Sp. Nov. Phylogenetics Sympatric Distribution Yunnan
资助者	Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015).
收录类别	SCI
语种	英语
资助者	Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015). ; Many thanks are owed to help from Jun-xiao Yang, Zi-jie Zhang, Wei-wei Zhou, Wang-sheng Ji, Shi-yang Dong, Vo Dina Ba, Nikolai Orlov, Ngoc Sang Nguyen, and Tran Ngoc Ninh in the field. We thank the local conservation departments for their help in field work of Thailand and Vietnam. The Institute of Animal for Scientific Purposes Development (IAD) issued permission (No.U1-01205-2558) in Thailand. The Ministry of Agriculture issued permission (No. 982/TCLN-BTTN) in Vietnam. We thank Liang Fei, Jodi J.L. Rowley and two anonymous reviewers for the invaluable comments on the manuscript. This work was supported by the program of Chinese Academy of Sciences (2015CASEABRI002), the National Natural Science Foundation of China (31090250), the Ministry of Science and Technology of China (MOST 2011FY120200), and the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences (the Large Research Infrastructure Funding). Chatmongkon Suwannapoom was sponsored by the Chinese Academy of Sciences Visiting Fellowship for Researchers (Postdoc) from Developing Countries (2013FFS130015).
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/10289
专题	科研部门_两栖爬行类多样性与进化（车静）遗传资源与进化国家重点实验室
通讯作者	chej@mail.kiz.ac.cn
作者单位	1.State Key Laboratory of Genetic Resources and Evolution State, Kunming Institute of Zoology, Chinese Academy of Sciences, Kun- ming 650223, Yunnan, China 2.Division of Fishery, School of Agriculture and Natural Resources, University of Phayao, Phayao, 56000, Thailand 3.Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar 4.Laboratory for Conservation and Utilization of Bio-resources, Yunnan University, Kunming 650091, China 5.Institute of Continuing Education, Sichuan Normal University, Chengdu 610068, Sichuan, China 6.School of Life Science, Henan University, Kaifeng 475001, Henan, China 7.College of life Sciences, Hainan Normal University, Haikou 570100, Hainan, China 8.Institute of Ecology and Biological Resources Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam 9.Centre for Biodiversity and Conservation Biology, Royal Ontario Museum, 100 Queen’s Park, Toronto M5S 2C6, Canada 10.University of Kansas Biodiversity Institute, 1345 Jayhawk Boulevard, Lawrence, Kansas 66045-7561, USA 11.Biology Department, James Madison University, 951 Carrier Dr., Harrisonburg, VA 22807 USA
推荐引用方式 GB/T 7714	Suwannapoom C,Yuan ZY,Wang LJ,et al. Taxonomic revision of the Chinese Limnonectes (Anura, Dicroglossidae) with the description of a new species from China and Myanmar.[J]. ZOOTAXA,2016,4093(2):181-200.
APA	Suwannapoom C.,Yuan ZY.,Wang LJ.,Nguyen TS.,Nguyen TQ.,...&chej@mail.kiz.ac.cn.(2016).Taxonomic revision of the Chinese Limnonectes (Anura, Dicroglossidae) with the description of a new species from China and Myanmar..ZOOTAXA,4093(2),181-200.
MLA	Suwannapoom C,et al."Taxonomic revision of the Chinese Limnonectes (Anura, Dicroglossidae) with the description of a new species from China and Myanmar.".ZOOTAXA 4093.2(2016):181-200.

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