KIZ OpenIR  > 实验与理论生态学
Host-Parasite Interactions and Purifying Selection in a Microsporidian Parasite of Honey Bees
Huang Q*1,2; Chen YP2; Wang RW*1,3; Cheng S4; Evans JD*2; qiang-huang@live.com; Jay.Evans@ars.usda.gov
2016
发表期刊PLOS ONE
卷号11期号:2页码:e0147549
摘要To clarify the mechanisms of Nosema ceranae parasitism, we deep-sequenced both honey bee host and parasite mRNAs throughout a complete 6-day infection cycle. By time-series analysis, 1122 parasite genes were significantly differently expressed during the reproduction cycle, clustering into 4 expression patterns. We found reactive mitochondrial oxygen species modulator 1 of the host to be significantly down regulated during the entire infection period. Our data support the hypothesis that apoptosis of honey bee cells was suppressed during infection. We further analyzed genome-wide genetic diversity of this parasite by comparing samples collected from the same site in 2007 and 2013. The number of SNP positions per gene and the proportion of non-synonymous substitutions per gene were significantly reduced over this time period, suggesting purifying selection on the parasite genome and supporting the hypothesis that a subset of N. ceranae strains might be dominating infection.
资助者This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784.
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资助者This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784. ; This study was supported by the United States Department of Agriculture—National Institute of Food and Agriculture- 2014-67013-21784.
文献类型期刊论文
条目标识符http://ir.kiz.ac.cn/handle/152453/10319
专题实验与理论生态学
遗传资源与进化国家重点实验室
通讯作者qiang-huang@live.com; Jay.Evans@ars.usda.gov
作者单位1.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Science, Kunming, 650223, China
2.USDA-ARS Bee Research Laboratory, BARC-East Building 306, Beltsville, Maryland, 20705, United States of America
3.Center for Ecological and Environmental Science, Northwestern Polytechnical University, Xi’an, 710072, China
4.Chongqing Academy of Animal Science, Chongqing, 402460, China
推荐引用方式
GB/T 7714
Huang Q*,Chen YP,Wang RW*,et al. Host-Parasite Interactions and Purifying Selection in a Microsporidian Parasite of Honey Bees[J]. PLOS ONE,2016,11(2):e0147549.
APA Huang Q*.,Chen YP.,Wang RW*.,Cheng S.,Evans JD*.,...&Jay.Evans@ars.usda.gov.(2016).Host-Parasite Interactions and Purifying Selection in a Microsporidian Parasite of Honey Bees.PLOS ONE,11(2),e0147549.
MLA Huang Q*,et al."Host-Parasite Interactions and Purifying Selection in a Microsporidian Parasite of Honey Bees".PLOS ONE 11.2(2016):e0147549.
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