KIZ OpenIR  > 基因起源组
Reciprocal genomic evolution in the ant-fungus agricultural symbiosis
Nygaard S*1; Hu HF2; Yang ZK2; Xie QL2; Ma CY2; Deng Y2; Dikow RB3; Rabeling C4,5; Li C2; Nash DR1; Wcislo WT6; Brady SG5; Schultz TR5; Zhang GJ*1,2,7; Boomsma JJ*1; Schiøtt M1; Chen ZS2; sanne@bio.ku.dk; guojie.zhang@bio.ku.dk; jjboomsma@bio.ku.dk
2016
发表期刊NATURE COMMUNICATIONS
卷号7期号:X页码:e12233
合作性质其它
摘要The attine ant-fungus agricultural symbiosis evolved over tens of millions of years, producing complex societies with industrial-scale farming analogous to that of humans. Here we document reciprocal shifts in the genomes and transcriptomes of seven fungus-farming ant species and their fungal cultivars. We show that ant subsistence farming probably originated in the early Tertiary (55-60 MYA), followed by further transitions to the farming of fully domesticated cultivars and leaf-cutting, both arising earlier than previously estimated. Evolutionary modifications in the ants include unprecedented rates of genome-wide structural rearrangement, early loss of arginine biosynthesis and positive selection on chitinase pathways. Modifications of fungal cultivars include loss of a key ligninase domain, changes in chitin synthesis and a reduction in carbohydrate-degrading enzymes as the ants gradually transitioned to functional herbivory. In contrast to human farming, increasing dependence on a single cultivar lineage appears to have been essential to the origin of industrial-scale ant agriculture.
资助者This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama.
收录类别SCI
语种英语
资助者This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama. ; This study was supported by grants from the Danish National Research Foundation (DNRF57) and the European Research Council (ERC Advanced grant 323085) (J.J.B.), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB13000000) (G.Z.), the Smithsonian Office of the Under Secretary for Science (T.R.S., S.G.B. and W.T.W.), the US National Science Foundation DEB-1456964 (C.R. and T.R.S.) and DEB-0949689 (T.R.S.), the US National Museum of Natural History (T.R.S.) and a University of Maryland-Smithsonian Institution Seed Grant (S.G.B. and T.R.S.). The Visitor’s Office of the Smithsonian Tropical Research Institute provided logistic help and facilities to work in Gamboa and the Autoridad Nacional del Ambiente y el Mar (ANAM) gave permission to sample and export ants from Panama.
文献类型期刊论文
条目标识符http://ir.kiz.ac.cn/handle/152453/10342
专题基因起源组
遗传资源与进化国家重点实验室
通讯作者sanne@bio.ku.dk; guojie.zhang@bio.ku.dk; jjboomsma@bio.ku.dk
作者单位1.Centre for Social Evolution, Department of Biology, University of Copenhagen, Universitetsparken 15, DK-2100 Copenhagen, Denmark
2.China National Genbank, BGI-Shenzhen, Shenzhen 518083, China
3.Smithsonian Institute for Biodiversity Genomics, Smithsonian Institution, Washington DC 20013-7012, USA
4.Department of Biology, University of Rochester, Rochester, New York 14627, USA.
5.Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013-7012, USA
6.Smithsonian Tropical Research Institute, Balboa, Anco ´n 03092, Panama
7.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, Chin
推荐引用方式
GB/T 7714
Nygaard S*,Hu HF,Yang ZK,et al. Reciprocal genomic evolution in the ant-fungus agricultural symbiosis[J]. NATURE COMMUNICATIONS,2016,7(X):e12233.
APA Nygaard S*.,Hu HF.,Yang ZK.,Xie QL.,Ma CY.,...&jjboomsma@bio.ku.dk.(2016).Reciprocal genomic evolution in the ant-fungus agricultural symbiosis.NATURE COMMUNICATIONS,7(X),e12233.
MLA Nygaard S*,et al."Reciprocal genomic evolution in the ant-fungus agricultural symbiosis".NATURE COMMUNICATIONS 7.X(2016):e12233.
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