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	Opportunities for biodiversity gains under the world's largest reforestation programme
	Hua FY*1; Wang XY2,3; Zhu JG2; Tang Y4; Yu DW*2,6; Wilcove DS*1,7; Zheng XL4; Fisher B5; Wang L2; hua.fangyuan@gmail.com; dougwyu@gmail.com; dwilcove@princeton.edu
	2016
发表期刊	NATURE COMMUNICATIONS
卷号	7期号:X页码:e12717
合作性质	其它
摘要	Reforestation is a critical means of addressing the environmental and social problems of deforestation. China's Grain-for-Green Program (GFGP) is the world's largest reforestation scheme. Here we provide the first nationwide assessment of the tree composition of GFGP forests and the first combined ecological and economic study aimed at understanding GFGP's biodiversity implications. Across China, GFGP forests are overwhelmingly monocultures or compositionally simple mixed forests. Focusing on birds and bees in Sichuan Province, we find that GFGP reforestation results in modest gains (via mixed forest) and losses (via monocultures) of bird diversity, along with major losses of bee diversity. Moreover, all current modes of GFGP reforestation fall short of restoring biodiversity to levels approximating native forests. However, even within existing modes of reforestation, GFGP can achieve greater biodiversity gains by promoting mixed forests over monocultures; doing so is unlikely to entail major opportunity costs or pose unforeseen economic risks to households.
资助者	Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327).
收录类别	SCI
语种	英语
资助者	Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327). ; Funding for this study was provided by the High Meadows Foundation and the 111 Project of China (B08037). D.W.Y. was supported by the National Natural Science Foundation of China (31400470, 41661144002), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology (GREKF13-13, GREKF14-13, and GREKF16-09). J.Z. and L.W. were supported by the National Natural Science Foundation of China (31272327).
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/10447
专题	科研部门_生态学与环境保护中心（俞维理） 遗传资源与进化国家重点实验室
通讯作者	hua.fangyuan@gmail.com; dougwyu@gmail.com; dwilcove@princeton.edu
作者单位	1.Program in Science, Technology, and Environmental Policy, Woodrow Wilson School of Public and International Affairs, Princeton University, Princeton, NJ 08544, USA. 2.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China 3.Kunming College of Life Sciences, University of Chinese Academy of Sciences, Kunming, Yunnan 650223, China. 4.College of Architecture and Environment, Sichuan University, Chengdu, Sichuan 610000, China 5.Gund Institute for Ecological Economics, Rubenstein School of Environment and Natural Resources, University of Vermont, Burlington, VT 05405, USA 6.School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich, Norfolk NR47TJ, UK. 7.Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544, USA
推荐引用方式 GB/T 7714	Hua FY*,Wang XY,Zhu JG,et al. Opportunities for biodiversity gains under the world's largest reforestation programme[J]. NATURE COMMUNICATIONS,2016,7(X):e12717.
APA	Hua FY*.,Wang XY.,Zhu JG.,Tang Y.,Yu DW*.,...&dwilcove@princeton.edu.(2016).Opportunities for biodiversity gains under the world's largest reforestation programme.NATURE COMMUNICATIONS,7(X),e12717.
MLA	Hua FY*,et al."Opportunities for biodiversity gains under the world's largest reforestation programme".NATURE COMMUNICATIONS 7.X(2016):e12717.

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2016090810.pdf（1286KB）	期刊论文	出版稿	开放获取	CC BY-NC-SA	请求全文

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