Phylogenetic relationships of megophryid frogs of the genus Leptobrachium (Amphibia, Anura) as revealed by mtDNA gene sequences

	Phylogenetic relationships of megophryid frogs of the genus Leptobrachium (Amphibia, Anura) as revealed by mtDNA gene sequences
	Matsui M*1; Hamidy A 1,2; Shimada T 1,7; Ahmad N 8,9; Belabut DM 8,10; Murphy RW 3,4; Khonsue W 5; Yambun P 6; Jiang JP 11; fumi@zoo.zool.kyoto-u.ac.jp
	2010
发表期刊	MOLECULAR PHYLOGENETICS AND EVOLUTION
卷号	56 期号:1 页码:259-272
合作性质	其它
摘要	By investigating genealogical relationships, we estimated the phylogenetic history and biogeography in the megophryid genus Leptobrachium (sensu lato, including Vibrissaphora) from southern China, Indochina, Thailand and the Sundaland. The genealogical relationships among the 30 named and unnamed taxa were estimated using 2009 bp of sequences from the mitochondrial DNA genes 12S rRNA, tRNA(val), and 16S rRNA using maximum parsimony, maximum likelihood, and Bayesian inference methods. The genus Leptobrachium was a well-supported monophyletic group that contained two major clades. One clade had three subclades primarily from disjunct regions including Borneo, Peninsular Malaysia and Java, and Thailand. The Bornean subclade included one species each from the Philippines and Sumatra. The other major clade consisted of two subclades, one from Indochina and the other from southern China (Vibrissaphora). Divergence times estimated an old evolutionary history of each subclade, one that could not be explained by the geohistory of Southeast Asian major landmasses.
关键词	Leptobrachium Sundaland Indochina Southern China Mtdna Phylogenetics
资助者	Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences.
收录类别	SCI
语种	英语
资助者	Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences. ; Field trips by MM were made possible by grants from The Monbusho Interna- tional Scientific Research Program (Field Research, 01041051, 02041051, 04041068, 06041066, and 08041144), The Mon- bukagakusho through the Japanese Society for the Promotion of Sciences (JSPS: Field Research, 15370038, 20405013), UKM (OUP- PLW-14-59/2008), and TJTTP-OECF. AH thanks M.D. Kusrini, S. Kir- ono, G.B. Mandigani, D. Susanto, M. Harvey, E. Smith, K.L. Sanders, T.Q. Nguyen, D.A. Anggraeni, and J. Rhamadani for providing tissue samples and to the Monbukagakusho for scholarship funding. RWM Funding for fieldwork was obtained through grants from the Natural Sciences and Engineering Research Council, Canada (Discovery Grant A3148), the Royal Ontario Museum (ROM) Foun- dation, and the ROM Volunteers Fieldwork Committee. All field- work was conducted using approved Animal Use Protocols. Fieldwork in Vietnam was assisted by N.L. Orlov, A. Lathrop, R.H. Bain, V.S. Cao, T.C. Ho and V.S. Nguyen; export permits were ob- tained with the assistance of V.S. Cao and the Institute of Ecology and Biological Resources, Hanoi. Specimens in Hainan Island were obtained while assisting H.-T. Shi, Hainan Normal University, and those from Yunnan were obtained with the assistance of D.-Q. Rao, Kunming Institute of Zoology and N.L. Orlov; research export permits were obtained through the Chinese Academy of Sciences.
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/10457
专题	科研部门_分子进化与基因组多样性（张亚平）遗传资源与进化国家重点实验室
通讯作者	fumi@zoo.zool.kyoto-u.ac.jp
作者单位	1.Graduate School of Human and Environmental Studies, Kyoto University, Kyoto 606-8501, Japan 2.Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Sciences, Gd. Widyasatwaloka, Jl. Raya Jakarta Bogor km 46, Cibinong West Java, Indonesia 3.Centre for Biodiversity and Conservation Biology, Royal Ontario Museum, 100 Queen’s Park, Toronto, Ont., Canada M5S 2C6 4.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, The Chinese Academy of Sciences, Kunming 650223, PR China 5.Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand 6.Research and Education Division, Sabah Parks, P.O. Box 10626, Kota Kinabalu 88806, Sabah, Malaysia 7.Faculty of Bioenvironmental Science, Kyoto Gakuen University, Kameoka, Kyoto 621-8555, Japan 8.Faculty of Science and Technology, Universiti Kebangsaan Malaysia, 43600 UKM Bangi, Selangor, Malaysia 9.Institute for Environment and Development (LESTARI), Universiti Kebangsaan Malaysia, 43600 UKM Bangi, Selangor, Malaysia 10.Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603 Kuala Lumpur, Malaysia 11.Chengdu Institute of Biology, The Chinese Academy of Sciences, Chengdu 610041, PR China
推荐引用方式 GB/T 7714	Matsui M*,Hamidy A,Shimada T,et al. Phylogenetic relationships of megophryid frogs of the genus Leptobrachium (Amphibia, Anura) as revealed by mtDNA gene sequences[J]. MOLECULAR PHYLOGENETICS AND EVOLUTION,2010,56(1):259-272.
APA	Matsui M*.,Hamidy A.,Shimada T.,Ahmad N.,Belabut DM.,...&fumi@zoo.zool.kyoto-u.ac.jp.(2010).Phylogenetic relationships of megophryid frogs of the genus Leptobrachium (Amphibia, Anura) as revealed by mtDNA gene sequences.MOLECULAR PHYLOGENETICS AND EVOLUTION,56(1),259-272.
MLA	Matsui M*,et al."Phylogenetic relationships of megophryid frogs of the genus Leptobrachium (Amphibia, Anura) as revealed by mtDNA gene sequences".MOLECULAR PHYLOGENETICS AND EVOLUTION 56.1(2010):259-272.

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