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	Identification and association of the single nucleotide polymorphisms in calpain3 (CAPN3) gene with carcass traits in chickens
	Zhang ZR1; Liu YP*1; Zhu Q1; Yao YG2; Jiang XS3; Du HR4; liuyp578@yahoo.com
	2009
发表期刊	BMC GENETICS
卷号	10期号:X页码:e10
合作性质	其它
摘要	Background: The aim of this study is to screen single nucleotide polymorphisms (SNP) of chicken Calpain3 (CAPN3) gene and to analyze the potential association between CAPN3 gene polymorphisms and carcass traits in chickens. We screened CAPN3 single nucleotide polymorphisms (SNP) in 307 meat-type quality chicken from 5 commercial pure lines (S01, S02, S03, S05, and D99) and 4 native breeds from Guangdong Province (Huiyang Huxu chicken and Qingyuan Ma chicken) and Sichuan Province (Caoke chicken and Shandi Black-bone chicken), China.  Results: Two SNPs (11818T > A and 12814T > G) were detected by single strand conformation polymorphism (SSCP) method and were verified by DNA sequencing. Association analysis showed that the 12814T > G genotypes were significantly associated with body weight (BW), carcass weight (CW), breast muscle weight (BMW), and leg muscle weight (LMW). Haplotypes constructed on the two SNPs (H1, TG; H2, TT; H3, AG; and H4, AT) were associated with BW, CW (P < 0.05), eviscerated percentage (EP), semi-eviscerated percentage (SEP), breast muscle percentage (BMP), and leg muscle percentage (LMP) (P < 0.01). Diplotype H1H2 was dominant for BW, CW, and LMP, and H2H2 was dominant for EP, SEP, and BMP.  Conclusion: We speculated that the CAPN3 gene was a major gene affecting chicken muscle growth and carcass traits or it was linked with the major gene(s). Diplotypes H1H2 and H2H2 might be advantageous for carcass traits.
资助者	This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences.
收录类别	SCI
语种	英语
资助者	This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences. ; This work was financially supported by Chinese national key technologies R & D programme (Grant No: 2008BADB2B01), the National 863 Project of China (No. 2008AA101001) and the provincial Eleventh five-year program on animal breeding of Sichuan Province (2006YZGG-19; 2006YZGG-25). YGY was supported by the "Century Program" (or Hundreds-Talent Pro- gram) of the Chinese Academy of Sciences.
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/10482
专题	科研部门_疾病机理遗传学和进化医学学科组（姚永刚） 科研部门_动物模型与人类重大疾病机理重点实验室
通讯作者	liuyp578@yahoo.com
作者单位	1.College of Animal Science and Technology, Sichuan Agriculture University, Ya'an, Sichuan 625014, PR China 2.Key Laboratory of Animal Models and Human Disease Mechanisms, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, PR China 3.Institute of Poultry Sciences, Sichuan Animal Science Academy, Chengdu, Sichuan 610066, PR China 4.Breeding Center, Sichuan Dahen Poultry Breeding Company, Chengdu, Sichuan 610066, PR China
推荐引用方式 GB/T 7714	Zhang ZR,Liu YP*,Zhu Q,et al. Identification and association of the single nucleotide polymorphisms in calpain3 (CAPN3) gene with carcass traits in chickens[J]. BMC GENETICS,2009,10(X):e10.
APA	Zhang ZR.,Liu YP*.,Zhu Q.,Yao YG.,Jiang XS.,...&liuyp578@yahoo.com.(2009).Identification and association of the single nucleotide polymorphisms in calpain3 (CAPN3) gene with carcass traits in chickens.BMC GENETICS,10(X),e10.
MLA	Zhang ZR,et al."Identification and association of the single nucleotide polymorphisms in calpain3 (CAPN3) gene with carcass traits in chickens".BMC GENETICS 10.X(2009):e10.

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