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	An integrative approach to reappraising species validity in Pseudexostoma (Teleostei: Sisoridae)
	Yang KF1,2; Jiang WS*1; Chen XY1,3; Zhou W4; Yang JX*1; jiangws@mail.kiz.ac.cn, yangjx@mail.kiz.ac.cn
	2016
发表期刊	ZOOTAXA
卷号	4158期号:3页码:352-366
摘要	The catfish genus Pseudexostoma (Sisoridae) currently contains three species: P. yunnanensis from the upper Irrawaddy River (Dayingjiang), and P. brachysoma and P. longipterus from the upper Salween River (Nujiang). We adopted an integrative approach combining both morphology-based (traditional measurements, truss-based and geometric morphometrics, fin skeleton examination and X-ray photography) and molecular approaches (genetic distances, haplotype network, phylogenetic relationships) to reappraise species validity in Pseudexostoma. Our results showed that P. longipterus cannot be distinguished from P. brachysoma from either morphological or molecular analyses, and that the former name should thus be considered a junior synonym of the latter. The two remaining geographically isolated species, P. yunnanensis and P. brachysoma, showed a distinct genetic distance equated to the typical 2% sequence divergence threshold delineating different species of fishes, and with a divergence time dated to approximately 1.22-2.14 million years ago. Although P. yunnanensis and P. brachysoma share some morphological characters that appear gradually diverging (body depth at anus and interorbital distance), some degree of overlap still exists, we suggest the two species are distinct based on the total evidence.
关键词	Glyptosternines Geometric Morphometrics X-ray Mitochondrial Dna Yunnan
资助者	This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences).
收录类别	SCI
语种	英语
资助者	This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences). ; This work was supported by the National Natural Science Foundation of China (31372182), Yunnan Provincial Science and Technology Program (2013FB070), the Chinese Academy of Sciences (KFJ-EW-STS-020), and U.S. NSF (Gaoligong Shan Biological Survey, DEB-0103795, to J. Slowinski and P. Fritsch, California Academy of Sciences).
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/10763
专题	科研部门_系统进化与生物地理学（杨君兴） 遗传资源与进化国家重点实验室 科研部门_东南亚野生动物多样性（陈小勇）
通讯作者	jiangws@mail.kiz.ac.cn, yangjx@mail.kiz.ac.cn
作者单位	1.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China 2.University of Chinese Academy of Sciences, Beijing 100049, China 3.Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar 4.Key Laboratory of Forest Disaster Warning & Control in Yunnan Province, Faculty of Forestry, Southwest Forestry University, Kun- ming, Yunnan 650224, China
推荐引用方式 GB/T 7714	Yang KF,Jiang WS*,Chen XY,et al. An integrative approach to reappraising species validity in Pseudexostoma (Teleostei: Sisoridae)[J]. ZOOTAXA,2016,4158(3):352-366.
APA	Yang KF,Jiang WS*,Chen XY,Zhou W,Yang JX*,&jiangws@mail.kiz.ac.cn, yangjx@mail.kiz.ac.cn.(2016).An integrative approach to reappraising species validity in Pseudexostoma (Teleostei: Sisoridae).ZOOTAXA,4158(3),352-366.
MLA	Yang KF,et al."An integrative approach to reappraising species validity in Pseudexostoma (Teleostei: Sisoridae)".ZOOTAXA 4158.3(2016):352-366.

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