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KIZ OpenIR  > 科研部门  > 转化基因组（李 明）
Matrine derivative YF-18 inhibits lung cancer cell proliferation and migration through down-regulating Skp2
Wu LC*1; Wang GZ2; Yang FF1,2; Li M5; Zhou GB2; Wang LS*1; Wei JR3; Huang N4; Zhang S1; wulichuan@126.com
2017
发表期刊Oncotarget
卷号8期号:7页码:11729-11738
摘要Lung cancer is the leading cause of cancer related death which needs novel drugs to improve patient outcomes. In this study, we examined the ability of YF-18, a novel matrine derivative to inhibit the growth and migration of lung cancer cells. By cell cycle analysis, wound healing and transwell assays, we found that YF-18 induced G2/M cell cycle arrest and inhibited migration of lung cancer cells in a dose-dependent manner. Further results indicated that YF-18 inhibited cell proliferation and migration through down-regulating Skp2 and up-regulating its substrates, p27 and E-cadherin. Moreover, YF-18 inhibited A549-luciferase cell xenograft tumor growth in a dose-dependent manner. The findings indicate that YF-18 bears therapeutic potentials for lung cancer.
关键词Skp2 Lung Cancer Matrine Derivative Migration Proliferation
资助者The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support.
收录类别SCI
语种英语
资助者The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support. ; The present study was financially supported by the National Natural Science Foundation of China (No. 21262005), the high level innovation team and outstanding scholar project of Guangxi institutions of higher education (gui jiao ren [2014] 49 hao), the Postdoctoral Science Foundation of Guangxi Province of China (Y201001928), the Scientific Research Foundation of Guangxi University (No. XJZ160924) the National Natural Science Funds for Distinguished Young Scholar (81425025), and the class General Financial Grant from the China Postdoctoral Science Foundation (2015M581172). We thank Professor Hu Tingjun of School of Zoology Science and Technology of Guangxi University for the experimental support.
文献类型期刊论文
条目标识符http://ir.kiz.ac.cn/handle/152453/10793
专题科研部门_转化基因组（李 明）
科研部门_动物模型与人类重大疾病机理重点实验室
通讯作者wulichuan@126.com
作者单位1.School of Chemistry and Chemical Engineering, Guangxi University, Nanning, Guangxi, PR China
2.State Key Laboratory of Membrane Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, PR China
3.Guangxi Scientific Research Center of Traditional Chinese Medicine, Guangxi University of Chinese Medicine, Nanning, Guangxi, PR China
4.Affiliated Tumour Hospital of Guangxi Medical University, Nanning, Guangxi, PR China
5.Key Laboratory of Animal Models and Human Disease Mechanisms of the Chinese Academy of Sciences and Yunnan Province, Kunming Institute of Zoology, Kunming, Yunnan, PR China
推荐引用方式
GB/T 7714		 Wu LC*,Wang GZ,Yang FF,et al. Matrine derivative YF-18 inhibits lung cancer cell proliferation and migration through down-regulating Skp2[J]. Oncotarget,2017,8(7):11729-11738.
APA Wu LC*.,Wang GZ.,Yang FF.,Li M.,Zhou GB.,...&wulichuan@126.com.(2017).Matrine derivative YF-18 inhibits lung cancer cell proliferation and migration through down-regulating Skp2.Oncotarget,8(7),11729-11738.
MLA Wu LC*,et al."Matrine derivative YF-18 inhibits lung cancer cell proliferation and migration through down-regulating Skp2".Oncotarget 8.7(2017):11729-11738.
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