Lead exposure through gestation-only caused long-term learning/memory deficits in young adult offspring

	Lead exposure through gestation-only caused long-term learning/memory deficits in young adult offspring
	Yang YX 1; Ma YM2; Zhang JY 2; Fan M 2; Liang CL 3; Cao J 1; Xu L1; Ni LX 2; Zhao SQ 2; Li LJ 2; lxu@vip.163.com
	2003
发表期刊	EXPERIMENTAL NEUROLOGY
卷号	184 期号:1 页码:489-495
摘要	Numerous observations in clinical and preclinical studies indicate that the developing brain is particular sensitive to lead (Pb)'s pernicious effects. However, the effect of gestation-only Pb exposure on cognitive functions at maturation has not been studied. We investigated the potential effects of three levels of Pb exposure (low, middle, and high Pb: 0.03%, 0.09%, and 0.27% of lead acetate-containing diets) at the gestational period on the spatial memory of young adult offspring by Morris water maze spatial learning and fixed location/visible platform tasks. Our results revealed that three levels of Pb exposure significantly impaired memory retrieval in male offspring, but only female offspring at low levels of Pb exposure showed impairment of memory retrieval. These impairments were not due to the gross disturbances in motor performance and in vision because these animals performed the fixed location/visible platform task as well as controls, indicating that the specific aspects of spatial learning/memory were impaired. These results suggest that exposure to Pb during the gestational period is sufficient to cause long-term learning/memory deficits in young adult offspring.
关键词	Hippocampus Water Maze Spatial Memory Gestation Lead Offspring Fetal Exposure
资助者	We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology.
收录类别	SCI
语种	英语
资助者	We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology. ; We wish to thank The National Basic Research Program G1999054000, National Natural Science Grants 39870280, 39925011, 39930080, Yunnan Science Grant 98C111M and the Research Program of The Chinese Academy of Sciences of China to L.X., and the Kunming Institute of Zoology.
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/10901
专题	科研部门_学习记忆的分子神经机制（徐林）
通讯作者	lxu@vip.163.com
作者单位	1.Laboratory of Learning and Memory, Kunming Institute of Zoology, The Chinese Academy of Sciences, Kunming 650223, Yunnan, PR China 2.Kunming Hospital of Children, Clinical Laboratory, Kunming 650034, Yunnan, PR China 3.Kunming General Hospital, Central Laboratory, Kunming, Yunnan, PR China
推荐引用方式 GB/T 7714	Yang YX,Ma YM*,Zhang JY,et al. Lead exposure through gestation-only caused long-term learning/memory deficits in young adult offspring[J]. EXPERIMENTAL NEUROLOGY,2003,184(1):489-495.
APA	Yang YX.,Ma YM*.,Zhang JY.,Fan M.,Liang CL.,...&lxu@vip.163.com.(2003).Lead exposure through gestation-only caused long-term learning/memory deficits in young adult offspring.EXPERIMENTAL NEUROLOGY,184(1),489-495.
MLA	Yang YX,et al."Lead exposure through gestation-only caused long-term learning/memory deficits in young adult offspring".EXPERIMENTAL NEUROLOGY 184.1(2003):489-495.

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