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	Phylogeny and systematic revision of the genus Typhlomys (Rodentia, Platacanthomyidae), with description of a new species
	Cheng F1,2; He K*2,3,4; Li JT6; Zhang BW*1,7; Jiang XL*2; Chen ZZ2,5; Zhang B2,5; Wan T2; zhangbw@ahu.edu.cn
	2017
发表期刊	Journal of Mammalogy
卷号	98期号:3页码:731-743
其他摘要	The “blind mouse” genus, Typhlomys, is an ancient taxon in the family Platacanthomyidae, comprising 2 living species and 4 living subspecies. We sequenced 3 mitochondrial (CytB, COI, and ND 2 ) and 2 nuclear (GHR and IRBP) gene segments to estimate the phylogenetic relationships of the species and subspecies, and to delimit species boundaries. We also evaluated patterns of morphological and morphometric variation of the skull. The molecular results suggest at least 5 species are present, corresponding to the 2 recognized species (Typhlomys cinereus and T. chapensis), 1 species previously considered a subspecies (T. ci. daloushanensis), 1 new species (Typhlomys nanus sp. nov.), and 1 putative new species, for which we lack adequate morphological specimens. We also determined that the subspecies Typhlomys cinereus jingdongensis is conspecific with T. chapensis. Diversification of the genus occurred in the late Miocene and Pliocene. The new species, T. nanus, is distributed in eastern and northeastern Yunnan at middle to high elevations (2,000 m to > 3,000 m above sea level). Interestingly, we detected a frameshift mutation within the IRBP gene, which is the 1st molecular evidence for the degradation of vision.
关键词	Frameshift Mutation Interphotoreceptor Retinoid-binding Protein (Irbp) Platacanthomyidae Species Delimitation Typhlomys
资助者	This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08).
收录类别	SCI
资助者	This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08). ; This study was supported by the National Nature Science Foundation of China (31272276), Yunnan Applied Basic Research Projects (2014FB176) and the fund of State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF15-08).
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/11022
专题	遗传资源与进化国家重点实验室 科研部门_兽类生态与进化（蒋学龙）
通讯作者	zhangbw@ahu.edu.cn
作者单位	1.Institute of Health Sciences, Anhui University, Hefei, Anhui 230601, China 2.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650204, China 3.Smithsonian Institution, National Museum of Natural History, 1000 Constitution Avenue, NW, Washington, DC 20004, USA 4.The Kyoto University Museum, Kyoto University, Kyoto 606-8501, Japan 5.Kunming College of Life Science, University of Chinese Academy of Sciences, Kunming 650204, China 6.Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China 7.School of Life Sciences, Anhui University, Hefei, Anhui 230601, China
推荐引用方式 GB/T 7714	Cheng F,He K*,Li JT,et al. Phylogeny and systematic revision of the genus Typhlomys (Rodentia, Platacanthomyidae), with description of a new species[J]. Journal of Mammalogy,2017,98(3):731-743.
APA	Cheng F.,He K*.,Li JT.,Zhang BW*.,Jiang XL*.,...&zhangbw@ahu.edu.cn.(2017).Phylogeny and systematic revision of the genus Typhlomys (Rodentia, Platacanthomyidae), with description of a new species.Journal of Mammalogy,98(3),731-743.
MLA	Cheng F,et al."Phylogeny and systematic revision of the genus Typhlomys (Rodentia, Platacanthomyidae), with description of a new species".Journal of Mammalogy 98.3(2017):731-743.

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