动物模型与人类重大疾病机理重点实验室知识库

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	Tree shrew (Tupaia belangeri) as a novel non-human primate laboratory disease animal model
	Xiao J1,2; Liu R2; Chen CS*2; chenc@mail.kiz.ac.cn
	2017
发表期刊	Zoological Research
卷号	38期号:3页码:127-137
摘要	The tree shrew (Tupaia belangeri) is a promising laboratory animal that possesses a closer genetic relationship to primates than to rodents. In addition, advantages such as small size, easy breeding, and rapid reproduction make the tree shrew an ideal subject for the study of human disease. Numerous tree shrew disease models have been generated in biological and medical studies in recent years. Here we summarize current tree shrew disease models, including models of infectious diseases, cancers, depressive disorders, drug addiction, myopia, metabolic diseases, and immune-related diseases. With the success of tree shrew transgenic technology, this species will be increasingly used in biological and medical studies in the future.
关键词	Tree Shrew (Tupaia Belangeri) Animal Model Transgenic Disease
资助者	This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222)
收录类别	其他
资助者	This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222) ; This study was supported by the National Nature Science Foundation of China (81325016, U1602221, 81322038 and U1502222)
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/11734
专题	科研部门_动物模型与人类重大疾病机理重点实验室 科研部门_肿瘤生物学（陈策实）
通讯作者	chenc@mail.kiz.ac.cn
作者单位	1.Medical Faculty of Kunming University of Science and Technology, Kunming Yunnan 650500, China 2.Key Laboratory of Animal Models and Human Disease Mechanisms of the Chinese Academy of Sciences and Yunnan Province, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming Yunnan 650223, China
推荐引用方式 GB/T 7714	Xiao J,Liu R,Chen CS*,et al. Tree shrew (Tupaia belangeri) as a novel non-human primate laboratory disease animal model[J]. Zoological Research,2017,38(3):127-137.
APA	Xiao J,Liu R,Chen CS*,&chenc@mail.kiz.ac.cn.(2017).Tree shrew (Tupaia belangeri) as a novel non-human primate laboratory disease animal model.Zoological Research,38(3),127-137.
MLA	Xiao J,et al."Tree shrew (Tupaia belangeri) as a novel non-human primate laboratory disease animal model".Zoological Research 38.3(2017):127-137.

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