动物模型与人类重大疾病机理重点实验室知识库

QQ客服 官方微博 反馈留言

	Atg5- and Atg7-dependent autophagy in dopaminergic neurons regulates cellular and behavioral responses to morphine
	Su LY1,2; Luo RC1,2; Ding YQ5; Xu L*1,4; Gao YG*1,2,3,4; Liu QJ1,2; Su JR1,2; Yang LX1; yaoyg@mail.kiz.ac.cn
	2017
发表期刊	Autophagy
卷号	**期号:**页码:Epub ahead of print
摘要	The molecular basis of chronic morphine exposure remains unknown. In this study, we hypothesized that macroautophagy/autophagy of dopaminergic neurons would mediate the alterations of neuronal dendritic morphology and behavioral responses induced by morphine. Chronic morphine exposure caused Atg5 (autophagy related 5)- and Atg7 (autophagy related 7)-dependent and dopaminergic neuron-specific autophagy resulting in decreased neuron dendritic spines and the onset of addictive behaviors. In cultured primary midbrain neurons, morphine treatment significantly reduced total dendritic length and complexity, and this effect could be reversed by knockdown of Atg5 or Atg7. Mice deficient for Atg5 or Atg7 specifically in the dopaminergic neurons were less sensitive to developing a morphine reward response, behavioral sensitization, analgesic tolerance and physical dependence compared to wild-type mice. Taken together, our findings suggested that the Atg5- and Atg7-dependent autophagy of dopaminergic neurons contributed to cellular and behavioral responses to morphine and may have implications for the future treatment of drug addiction.
关键词	Atg5 Atg7 Addiction Autophagy Dopaminergic Neuron Morphine
资助者	The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005).
收录类别	SCI
语种	英语
资助者	The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005). ; The study was done at Kunming Institute of Zoology, Chinese Academy of Sciences (CAS), and was supported by the National Natural Science Foundation of China (31671050), the Strategic Priority Research Program (B) of CAS (XDB02020003) and the Bureau of Frontier Sciences and Education of CAS (QYZDJ-SSW-SMC005).
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/11946
专题	科研部门_动物模型与人类重大疾病机理重点实验室 科研部门_学习记忆的分子神经机制（徐林） 科研部门_疾病机理遗传学和进化医学学科组（姚永刚） 中国科学院昆明灵长类研究中心
通讯作者	yaoyg@mail.kiz.ac.cn
作者单位	1.Key Laboratory of Animal Models and Human Disease Mechanisms of the Chinese Academy of Sciences & Yunnan Province, Kunming Institute of Zoology, Kunming, Yunnan, China 2.Kunming College of Life Science,University of Chinese Academy of Sciences, Kunming, Yunnan, China 3.Kunming Primate Research Center of the Chinese Academy of Sciences, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China 4.CAS Center for Excellence in Brain Science and Intelligence Technology, Chinese Academy of Sciences, Shanghai, China 5.Key Laboratory of Arrhythmias, Department of Anatomy and Neurobiology, Tongji University School of Medicine Shanghai, Shanghai, China
推荐引用方式 GB/T 7714	Su LY,Luo RC,Ding YQ,et al. Atg5- and Atg7-dependent autophagy in dopaminergic neurons regulates cellular and behavioral responses to morphine[J]. Autophagy,2017,**(**):Epub ahead of print.
APA	Su LY.,Luo RC.,Ding YQ.,Xu L*.,Gao YG*.,...&yaoyg@mail.kiz.ac.cn.(2017).Atg5- and Atg7-dependent autophagy in dopaminergic neurons regulates cellular and behavioral responses to morphine.Autophagy,**(**),Epub ahead of print.
MLA	Su LY,et al."Atg5- and Atg7-dependent autophagy in dopaminergic neurons regulates cellular and behavioral responses to morphine".Autophagy **.**(2017):Epub ahead of print.

条目包含的文件
文件名称/大小	文献类型	版本类型	开放类型	使用许可
2017072401.pdf（3361KB）	期刊论文	出版稿	开放获取	CC BY-NC-SA	请求全文

个性服务

推荐该条目

保存到收藏夹

查看访问统计

导出为Endnote文件

谷歌学术

谷歌学术中相似的文章

[Su LY]的文章

[Luo RC]的文章

[Ding YQ]的文章

百度学术

百度学术中相似的文章

[Su LY]的文章

[Luo RC]的文章

[Ding YQ]的文章

必应学术

必应学术中相似的文章

[Su LY]的文章

[Luo RC]的文章

[Ding YQ]的文章

相关权益政策

暂无数据

收藏/分享

除非特别说明，本系统中所有内容都受版权保护，并保留所有权利。