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	Taxonomic revision of the genus Mesechinus (Mammalia: Erinaceidae) with description of a new species
	Huai-Sen Ai; Kai He; Wei-Ting Su; Xue-Long Jiang; Zhong-Zheng Chen; Jia-Qi Li; Tao Wan; Quan Li; Wen-Hui Nie; Jin-Huan Wang; hekai@mail.kiz.ac.cn
	2018
发表期刊	ZOOLOGICAL RESEARCH
卷号	39期号:5页码:335–347
摘要	Hedgehogs in the genus Mesechinus (FamilyErinaceidae), which include two currently recognizedspecies (M. dauuricus and M. hughi), are distributedfrom northeast Mongolia to the upper Amur Basin inRussia and adjacent areas in northeast and northernChina. In recent years, a population of Mesechinushedgehogs was discovered from Mt. Gaoligong,southwestern Yunnan, China, far from the knowndistribution range of the genus. Furthermore, thesehedgehogs are the only known population to bedistributed at elevations higher than 2 100 m and insympatry with gymnures. To evaluate the taxonomicstatus of these hedgehogs, we examined specimensrepresenting Mesechinus taxa in China and furtherconducted morphometric and karyotypic analyses.Our results supported the existence of four speciesin China. Specifically, we identified the hedgehogsfrom Mt. Gaoligong as a new species, Mesechinuswangi sp. nov. , and recognized M. miodon, previouslyconsidered as a synonym of either M. dauuricus or M.hughi, as a distinct species. Interestingly, we observeda supernumerary M 4 on all specimens of Mesechinuswangi sp. nov. , which is an extremely rare event inthe evolution of mammalian dentition.
关键词	Mesechinus Taxonomy Morphometrics Inhibitory Cascade Karyotype New Species Supernumerary Molar
资助者	This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092)
DOI	10.24272/j.issn.2095-8137.2018.034
收录类别	其他
语种	英语
资助者	This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092) ; This study was supported by the National Key Research and Development Program of China (2017YFC0505200), Fundamental Research Funds for NIES in 2017(GYZX170308) and Biodiversity Conservation Program by MEP of China, K. H. was supported by a JSPS Postdoctoral Fellowship for Overseas Researchers (P16092)
引用统计
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/12132
专题	科研部门_兽类生态与进化（蒋学龙）
通讯作者	hekai@mail.kiz.ac.cn
作者单位	1.Gaoligongshan National Nature Reserve, Baoshan Yunnan 678000, China 2.Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming Yunnan 650223, China 3.The Kyoto University Museum, Kyoto University, Kyoto 606-8501, Japan 4.College of Life Sciences, Anhui Normal University, Wuhu Anhui 241000, China 5.Nanjing Institute of Environmental Sciences, Ministry of Environmental Protection, Nanjing Jiangsu 210042, China
推荐引用方式 GB/T 7714	Huai-Sen Ai,Kai He,Wei-Ting Su,et al. Taxonomic revision of the genus Mesechinus (Mammalia: Erinaceidae) with description of a new species[J]. ZOOLOGICAL RESEARCH,2018,39(5):335–347.
APA	Huai-Sen Ai.,Kai He.,Wei-Ting Su.,Xue-Long Jiang.,Zhong-Zheng Chen.,...&hekai@mail.kiz.ac.cn.(2018).Taxonomic revision of the genus Mesechinus (Mammalia: Erinaceidae) with description of a new species.ZOOLOGICAL RESEARCH,39(5),335–347.
MLA	Huai-Sen Ai,et al."Taxonomic revision of the genus Mesechinus (Mammalia: Erinaceidae) with description of a new species".ZOOLOGICAL RESEARCH 39.5(2018):335–347.

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