| Antimicrobial peptide diversity in the skin of the torrent frog, Amolops jingdongensis | |
| He XQ1; Yang SL2; Wei L1; Liu R2; Lai R*1,2; Rong MQ*1,2; rlai72@njau.edu.cn; rongmingqiang@yahoo.com | |
| 2013 | |
| 发表期刊 | AMINO ACIDS
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| 卷号 | 44期号:2页码:481-487 |
| 合作性质 | 其它 |
| 摘要 | Antimicrobial peptide diversity has been found in some amphibians. The diversity of antimicrobial peptides may have resulted from the diversity of microorganisms encountered by amphibians. Peptidomics and genomics analyses were used to study antimicrobial peptide diversity in the skin secretions of the torrent frog, Amolops jingdongensis. Thirty-one antimicrobial peptides belonging to nine groups were identified in the skin secretions of this frog. Among them, there are two novel antimicrobial groups (jingdongin-1 and -2) with unique structural motifs. The other seven groups belong to known antimicrobial peptide families, namely brevinin-1, brevinin-2, odorranain-F, esculentin-2, temporin, amolopin-3, and ranacyclin. Combined with previous reports, more than 13 antimicrobial peptide groups have been identified from the genus Amolops. Most of these antimicrobial peptide groups are also found in amphibians belonging to the genus Rana or Odorrana which suggests a possible evolutionary connection among Amolops, Rana, and Odorrana. Two novel antimicrobial groups (jingdongin-1 and -2) were synthesized and their antimicrobial activities were assayed. Some of them showed strong antimicrobial abilities against microorganisms including Gram-negative and -positive bacteria, and fungi. The extreme diversity of antimicrobial peptides in the Amolops amphibians was demonstrated. In addition, several novel peptide templates were provided for antimicrobial agent design. |
| 关键词 | Antimicrobial Peptides Amphibian Skin Diversity Skin Amolops Amolops |
| 资助者 | This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. |
| 语种 | 英语 |
| 资助者 | This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. ; This work was supported by Chinese National Natural Science Foundation (30830021, 31025025, 31070701, 31000960, 31025025, U1132601), the Ministry of Science and Technology (2010CB529800, 2009ZX09103-1/091, 2011ZX09102- 002-10), the Ministry of Agriculture (2009ZX08009-159B) and Yunnan Province Y103951111. |
| WOS记录号 | WOS:000313794600018 |
| 引用统计 | |
| 文献类型 | 期刊论文 |
| 条目标识符 | http://ir.kiz.ac.cn/handle/152453/7223 |
| 专题 | 科研部门_天然药物功能蛋白质学科组（赖仞） 科研部门_动物模型与人类重大疾病机理重点实验室 |
| 通讯作者 | rlai72@njau.edu.cn; rongmingqiang@yahoo.com |
| 作者单位 | 1.Life Sciences College of Nanjing Agricultural University, Nanjing 210095, Jiangsu, China 2.Key Laboratory of Animal Models and Human Disease Mechanisms of the Chinese Academy of Sciences and Yunan Provinces, Kunming Institute of Zoology, Kunming 650223, Yunnan, China |
| 推荐引用方式 GB/T 7714 | He XQ,Yang SL,Wei L,et al. Antimicrobial peptide diversity in the skin of the torrent frog, Amolops jingdongensis[J]. AMINO ACIDS,2013,44(2):481-487. |
| APA | He XQ.,Yang SL.,Wei L.,Liu R.,Lai R*.,...&rongmingqiang@yahoo.com.(2013).Antimicrobial peptide diversity in the skin of the torrent frog, Amolops jingdongensis.AMINO ACIDS,44(2),481-487. |
| MLA | He XQ,et al."Antimicrobial peptide diversity in the skin of the torrent frog, Amolops jingdongensis".AMINO ACIDS 44.2(2013):481-487. |
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